In press

    Tabuce R., Lihoreau F., Mees F., Orliac M., De Putter T. & Smith T., in press. A reassessment of the Oligocene hyracoids from Malembo, Cabinda, Angola. Geobios. doi:
    Solé F., Morse P. E., Bloch J. I., Gingerich P. D. & Smith T., in press. On the evolution of the mesonychid Dissacus across the PETM event: to be, or not to be a “dwarfed” species? Geobios. doi:
    Solé F. & Morlo M., in press. New hyaenodonts (Mammalia) from the Ypresian locality of Prémontré (France) support a radiation of the hyaenodonts in Europe already at the end of the early Eocene. Geobios. doi:
    Rana R.S., Waqas M., Orliac M., Folie A. & Smith T., in press. A new basal raoellid artiodactyl (Mammalia) from the middle Eocene Subathu Group of Rajouri District, Jammu and Kashmir, northwest Himalaya, India. Geobios. doi:
    Folie A., Mees F., De Putter T. & Smith T., in press. Presence of the large aquatic snake Palaeophis africanus in the middle Eocene marine margin of the Congo Basin, Cabinda, Angola. Geobios. doi: 10.1016/j.geobios.2020.11.002.
    Pérez-García A. & Smith T., in press. Systematics and diversity of the giant soft-shelled turtles (Cryptodira, Trionychidae) from the earliest Eocene of Belgium. Geobios. doi: 10.1016/j.geobios.2020.07.006.
    Zaher H., Folie A., Quadros A.B., Rana R.S., Kumar K., Rose K.D., Fahmy M. & Smith T., in press. Additional vertebral material of Thaumastophis (Serpentes: Caenophidia) from the early Eocene of India provides new insights on the early diversification of colubroidean snakes. Geobios. doi: 10.1016/j.geobios.2020.06.009.


    Yans J., Verhaert M., Gautheron C., Antoine P.-O., Moussi B., Dekoninck A., Decrée S., Chaftar H.-R., Hatira N., Dupuis C., Pinna-Jamme R., Jamoussi F., 2021. (U-Th)/He dating of Supergene Iron (Oxyhydr-)Oxides of the Nefza-Sejnane District (Tunisia): New Insights into mineralization and mammalian biostratigraphy. Minerals 11(3): 260 (19 pages). doi: 10.3390/min11030260.
    Solé F., Fisher V., Denayer J., Speijer R., Fournier M., Le Verger K., Ladevèze S., Folie A. & Smith T., 2021. The upper Eocene-Oligocene carnivorous mammals from the Quercy Phosphorites (France) housed in Belgian collections. Geologica Belgica 24(1-2):1-16. doi: 10.20341/gb.2020.006.
    Zack S., Rose K.D., Holbrook L.T., Kumar K., Rana R.S. & Smith T., 2021. An enigmatic new ungulate-like mammal from the early Eocene of India. Papers in Paleontology 7: 497-520. doi: 10.1002/spp2.1288.


    Mayr G., Gingerich P.D. & Smith T., 2020. Skeleton of a new owl from the early Eocene of North America (Aves, Strigiformes) with an accipitrid-like foot morphology. Journal of Vertebrate Paleontology, 40(2):e1769116. doi: 10.1080/02724634.2020.1769116.
    Rose K.D., Holbrook L.T., Rana R.S., Ahrens H.E., Dunn R.H., Folie A., Jones K.E. & Smith T., 2020. Anatomy, Relationships, and Paleobiology of Cambaytherium (Mammalia, Perissodactylamorpha, Anthracobunia) from the lower Eocene of western India. Journal of Vertebrate Paleontology 39 (sup. 1): 1-147. doi: 10.1080/02724634.2020.1761370.
    De Putter T. & Ruffet G., 2020. Supergene manganese ore records 75 Myr-long Campanian to Pleistocene geodynamic evolution and weathering history of the Central African Great Lakes Region – Tectonics drives, climate assists. Gondwana Research, 83: 96-117. doi: 10.1016/ .
    Solé F., De Bast E., Legendre H., Rana S.R., Kumar K., Rose K.D. & Smith T., 2020. Chapter 12. New specimens of Frugivastodon (Mammalia: Apatotheria) from the early Eocene of India confirm its apatemyid status and elucidate dispersal of Apatemyidae. In: G. V. R. Prasad and R. Patnaik (eds.), Biological Consequences of Plate Tectonics: New Perspectives on Post-Gondwana Break-up—A Tribute to Ashok Sahni, Vertebrate Paleobiology and Paleoanthropology Series, Springer, pp.279-304. doi: 10.1007/978-3-030-49753-8_12.
    Steeman T., De Weirdt J., Smith T., De Putter T., Mees F. & Louwye S., 2020. Dinoflagellate cyst biostratigraphy and palaeoecology of the early Paleogene Landana reference section, Cabinda Province, Angola. Palynology 44(2): 280-309. doi: 10.1080/01916122.2019.1575091.
    Perez-Garcia A., Mees F. & Smith T., 2020. Shell anatomy of the African Paleocene bothremydid turtle Taphrosphys congolensis and systematic implications within Taphrosphyini. Historical Biology, 32(3): 376-385. doi: 10.1080/08912963.2018.1497023.


    Mayr G. & Smith T., 2019. New Paleocene bird fossils from the North Sea Basin in Belgium and France. Geologica Belgica, 22(1-2): 35-46. doi: 10.20341/gb.2019.003.
    Mayr G. & Smith T., 2019. A diverse bird assemblage from the Ypresian of Belgium furthers knowledge of Early Eocene avifaunas of the North Sea BasinN. Jb. Geol. Paläont. Abh., 291(3): 253-281. doi: 10.1127/njgpa/2019/0801.
    Solé F., Plateau O., Le Verger K. & Phélizon A., 2019. New paroxyclaenid mammals from the early Eocene of the Paris Basin (France) shed light on the origin and evolution of these endemic European cimolestans. Journal of Systematic Palaeontology, 17 (20): 1711-1743. doi: 10.1080/14772019.2018.1551248.
    Morse P.E., Chester S.G.B, Boyer D.M., Smith T., Smith R., Gigase P. & Bloch J.I., 2019. New fossils, systematics, and biogeography of the oldest known crown primate Teilhardina from the earliest Eocene of Asia, Europe, and North America. Journal of Human Evolution, 128: 103-131. doi: 10.1016/j.jhevol.2018.08.005.
    Solé F., Noiret C., Desmares D., Adnet S., Taverne L., De Putter T., Mees F., Yans J., Steeman T., Louwye S., Folie A., Stevens N.J., Gunnell G.F., Baudet D., Kitambala Yaya N. & Smith T., 2019. Reassessment of historical sections from the Paleogene margin of the Congo Basin reveals an almost complete absence of Danian deposits. Geoscience Frontiers, 10: 1039-1063. doi: 10.1016/j.gsf.2018.06.002.
    Mayr G., Gingerich P.D. & Smith T., 2019. Calcardea junnei Gingerich, 1987 from the late Paleocene of North America is not a heron, but resembles the early Eocene Indian taxon Vastanavis Mayr et al., 2007.Journal of Paleontology, 93(2): 359-367. doi: 10.1017/jpa.2018.85.
    Taverne L., De Putter T., Mees F. & Smith T., 2019. Cabindachanos dartevellei gen. and sp. nov., a new chanid fish (Ostariophysi, Gonorynchiformes) from the marine Paleocene of Cabinda (Central Africa). Geologica Belgica, 22(1-2): 1-6. doi: 10.20341/gb.2018.011.


    Koenigswald W. von, Rose, K.D., Holbrook L.T., Kumar K., Rana R.S. & Smith T., 2018. Mastication and enamel microstructure in Cambaytherium, a perissodactyl‑like ungulate from the early Eocene of India. Paläontologische Zeitschrift, 92:671–680. doi: 10.1007/s12542-018-0422-8.
    Rose K.D., Dunn R.H., Kumar K., Perry J.M.G., Prufrock, K.A., Rana, R.S. & Smith T., 2018. New fossils from Tadkeshwar Mine (Gujarat, India) increase primate diversity from the early Eocene Cambay Shale. Journal of Human Evolution, 122:93-107. doi: 10.1016/j.jhevol.2018.05.006.
    De Bast E., Gagnaison C. & Smith T., 2018. Plesiadapid mammals from the latest Paleocene of France offer new insights on the evolution of Plesiadapis during the Paleocene-Eocene transition. Journal of Vertebrate Paleontology, 38(3): e1460602. doi: 10.1080/02724634.2018.1460602.
    Müller J., Roberts E.M., Naylor E.R., Stevens N.J., 2018. A fossil gekkotan (Squamata) from the Late Oligocene Nsungwe Formation, Rukwa Rift Basin, Tanzania. Journal of Herpetology, 52(2):223-227. doi: 10.1670/17-123.
    Epa Y.R., Stigall A.L., O’Brien H.D., Roberts E.M. & Stevens N.J., 2018. Morphological diversification of ampullariid gastropods (Nsungwe Formation, Late Oligocene, Rukwa Rift Basin) is coincident with onset of East African Rifting. Papers in Palaeontology, 4(3): 327-348. doi: 10.1002/spp2.1108.
    Mayr G., De Pietri V., Scofield R.P. & Smith T., 2018. A fossil heron from the early Oligocene of Belgium: the earliest temporally well-constrained record of the Ardeidae. IBIS, 161:79-90. doi: 10.1111/ibi.12600.


    Benammi Mou., Adnet S., Marivaux L., Yans J., Noiret C., Tabuce R., Surault J., El Kati I., Enault S., Baidder L., Saddiqi O., Benammi Moh., 2017. Geology, biostratigraphy and carbon isotope chemostratigraphy of the Paleogene fossil-bearing Dakhla sections, Southwestern Moroccan Sahara. Geological Magazine, 156(1): 117-132. doi: 10.1017/S0016756817000851.
    Borths M. & Stevens NJ., 2017. The first African hyaenodont from the beginning of the Paleogene-Neogene carnivore transition (late Oligocene, Nsungwe Formation, Rukwa Rift Basin, Tanzania). PLOS ONE, 12(10): e0185301. doi: 10.1371/journal.pone.0185301.
    Bastl K., Nagel D. & Solé F., 2017. Incus facet morphology in carnivorous mammals from different ecosystems: taxonomy vs. habitat. Geobios, 16 : 284-302. doi: 10.1016/j.crpv.2016.11.008.
    Solé F. & Ladevèze S., 2017. Evolution of the hypercarnivorous dentition in mammals (Metatheria, Eutheria) and its bearings on the development of the tribosphenic molars. Evolution & Development, 19, 56-68. doi: 10.1111/ede.12219.
    Mayr G. & Smith T., 2017. First Old World record of the poorly known, swan-sized anseriform bird Paranyroca from the late Oligocene/early Miocene of France. Geologie und Paläontologie, 286(3): 349-354. doi: 10.1127/njgpa/2017/0703.
    Bemis K.E., Tyler J.C., Bemis W.E., Kumar K., Rana R.S. & Smith T., 2017. A gymnodont fish jaw with remarkable molariform teeth from the early Eocene of Gujarat, India (Teleostei, Tetraodontiformes). Journal of Vertebrate Paleontology, e1369422. doi: 10.1080/02724634.2017.1369422.
    Delfino M., Martin J.E., de Lapparent de Broin F. & Smith T., 2017. Evidence for a pre-PETM dispersal of the earliest European crocodyloids. Historical Biology, 31(7): 845-852. doi: 10.1080/08912963.2017.1396323.
    Perez-Garcia A. & Smith T., 2017. Identification of the African–European Erymnochelys group (Pleurodira, Podocnemididae) in the Belgian fossil record: first finding of Eocenochelus eremberti outside its type locality. Fossil Record, 20: 245-251. doi: 10.5194/fr-20-245-2017.
    Dagosto M., Gebo D., Ni X. & Smith T., 2017. Estimating body size in early primates: The case of Archicebus and Teilhardina. Journal of Human Evolution, 115: 8-19. doi: 10.1016/j.jhevol.2017.02.005.
    Gunnell G.F., Smith R. & Smith T., 2017. 33 million year ols Myotis (Chiroptera, Vespertilionidae) and the rapid global radiation of modern bats. Plos One, 12(3): e0172621. doi: 10.1371/journal.pone.0172621.
    Solé F., Godinot M., Laurent Y., Galoyer A. & Smith, T. 2017. The European mesonychid mammals : phylogeny, ecology, biogeography, and biochronology. Journal of Mammalian Evolution, 25: 339-379. doi: 10.1007/s10914-016-9371-8.
    Taverne L., 2017. New data on the osteoglossid fishes (Teleostei, Osteoglossiformes) from the marine Danian (Paleocene) of Landana (Cabinda Enclave, Angola). Geo-Eco-Trop, 40(4): 297-304. .
    Miller E.R., Gunnell G.F., Seiffert E.R., Sallam H.M. & Schwartz G.T., 2017. Patterns of dental emergence in early anthropoid primates from the Fayum Depression, Egypt. Historical Biology, 30(1-2): 157-165. doi: 10.1080/08912963.2017.1294169.


    De Bast E. & Smith, T., 2016. The oldest Cenozoic mammal fauna of Europe: implication of the Hainin reference fauna for mammalian evolution and dispersals during the Palaeocene. Journal of Systematic Palaeontology, 15(9): 741-785. doi: 10.1080/14772019.2016.1237582.
    Smith T., Kumar K., Rana R.S., Folie A., Solé F., Noiret C., Steeman T., Sahni A. & Rose K.D., 2016. New early Eocene vertebrate assemblage from western India reveals a mixed fauna of European and Gondwana affinities, Geoscience Frontiers, 7: 969-1001. doi: 10.1016/j.gsf.2016.05.001.
    Solé F., Essid E.M., Marzougui W., Temani R., Khayati Ammar H., Mahboubi M., Marivaux L., Vianey-Liaud M. & Tabuce R., 2016. New fossils of Hyaenodonta (Mammalia) from the Eocene localities of Chambi (Tunisia) and Bir el Ater (Algeria), and the evolution of the earliest African hyaenodonts. Palaeontologia Electronica, 19.2.38A: 1-23. doi: 10.26879/598.
    Sallam H.M., Sileem A.H., Miller E.R. & Gunnell G.F., 2016. Deciduous dentition and dental eruption sequence of Bothriogenys fraasi (Anthracotheriidae, Artiodactyla) from the Fayum Depression, Egypt. Palaeontologia Electronica, 19.3.38A: 1-17. doi: 10.26879/646.
    Dunn R.H., Rose K.D., Rana R.S., Kumar K., Sahni A. & Smith, T., 2016. New euprimate postcrania from the early Eocene of Gujarat, India, and the strepsirrhine-haplorhine divergence. Journal of Human Evolution, 99: 25-51. doi: 10.1016/j.jhevol.2016.06.006.
    Noiret C., Steurbaut E., Tabuce R., Marandat B., Schnyder J., Storme J-Y. & Yans J., 2016. New bio-chemostratigraphic dating of a unique early Eocene sequence from southern Europe results in precise mammalian biochronological tie-points. Newsletters on Stratigraphy, 49(3): 469-480. doi: 10.1127/nos/2016/0336.
    Pérez-García A., Smith R. & Smith T., 2016. Anatomy and variability of Cuvierichelys parisiensis, a geoemydid turtle that crosses the Eocene-Oligocene boundary in Belgium. Journal of Iberian Geology, 42(2): 161-170. doi: 10.5209/rev_JIGE.2016.v42.n2.51368.
    Solé F., Smith T., De Bast E., Codrea V. & Gheerbrant E., 2016. New carnivoraforms from the latest Paleocene of Europe and their bearing on the origin and radiation of Carnivoraformes (Carnivoramorpha, Mammalia). Journal of Vertebrate Paleontology, 36(2): e1082480. doi: 10.1080/02724634.2016.1082480.


    De Putter T., Ruffet G., Yans J. & Mees F., 2015. The age of supergene manganese deposits in Katanga and its implications for the Neogene evolution of the African Great Lakes Region. Ore Geology Reviews, 71: 350–362. doi: 10.1016/j.oregeorev.2015.06.015. .
    Solé F., Falconnet J. & Vidalenc D., 2015. New fossil Hyaenodonta (Mammalia, Placentalia) from the Ypresian and Lutetian of France and the evolution of the Proviverrinae in Southern France. Palaeontology, 58: 1049–1072. doi: 10.1111/pala.12198.
    Smith T., Solé F., Missiaen P., Rana R., Kumar K., Sahni A. & Rose K.D., 2015. First early Eocene tapiroid from India and its implication for the paleobiogeographic origin of perissodactyls. Palaeovertebrata, 39 (2): e5. doi: 10.18563/pv.39.2.e5.
    Solé F., Amson E., Borths M., Vidalenc D., Morlo M. & Bastl K., 2015. A New Large Hyainailourine from the Bartonian of Europe and Its Bearings on the Evolution and Ecology of Massive Hyaenodonts (Mammalia). PLoS ONE, 10(9): e0135698. doi: 10.1371/journal.pone.0135698.
    Rana R.S., Kumar K., Zack S.P., Solé F., Rose K.D., Missiaen P., Singh L., Sahni A. & Smith T., 2015. Craniodental and postcranial morphology of Indohyaenodon raoi from the early Eocene of India, and its implications for ecology, phylogeny, and biogeography of hyaenodontid mammals. Journal of Vertebrate Paleontology, 35(5): e965308. doi: 10.1080/02724634.2015.965308.
    Solé F., Smith T., Tabuce R. & Marandat B., 2015. New dental elements of the oldest proviverrine mammal from the early Eocene of Southern France support possible African origin of the subfamily. Acta Palaeontologica Polonica, 60(3): 527–538. doi: 10.4202/app.00146.2014.
    Blackburn D.C., Roberts E.M. & Stevens N.J., 2015. The earliest record of the endemic African frog family Ptychadenidae from the Oligocene Nsungwe Formation of Tanzania, Journal of Vertebrate Paleontology, 35:2, e907174. doi: 10.1080/02724634.2014.907174.
    Gebo D.L., Smith R., Dagosto M. & Smith T., 2015. Additional Postcranial Elements of Teilhardina belgica, the oldest European primate. American Journal of Physical Anthropology, 156: 388-406. doi: 10.1002/ajpa.22664.


    Martin J., Smith T., de Lapparent de Broin F., Escuillié F. & Delfino M., 2014. Late Palaeocene eusuchian remains from Mont de Berru, France, and the origin of the alligatoroid Diplocynodon. Zoological Journal of the Linnean Society, 172: 867-891. doi: 10.1111/zoj.12195.
    Rose K.D., Holbrook L.T., Rana R.S., Kumar K., Jones K.E., Ahrens H.E., Missiaen P., Sahni A.A. & Smith T., 2014. Early Eocene fossils suggest the mammalian order Perissodactyla originated in India. Nature Communications, 5: 5570. doi: 10.1038/ncomms6570.
    Storme J.-Y., Steurbaut E., Devleeschouwer X., Dupuis C., Iacumin P., Rochez G. & Yans J., 2014. Integrated bio-chemostratigraphical correlations and climatic evolution across the Danian-Selandian boundary at low latitudes. Palaeogeography, Palaeoclimatology, Palaeoecology, 414: 212-224. doi: 10.1016/j.palaeo.2014.09.002.
    Yans J., Marandat B., Masure E., Serra-Kiel J., Schnyder J., Storme J.-Y., Marivaux L., Adnet S., Vianey-Liaud M. & Tabuce R., 2014. Refined bio- (benthic foraminifera, dinoflagellate cysts) and chemostratigraphy (13Corg) of the earliest Eocene at Albas-Le Clot (Corbières, France): implications for mammalian biochronology in Western Europe. Newsletters on Stratigraphy, 47/3: 331-353. doi: 10.1127/nos/2014/0050.
    McCartney J., Stevens N.J. & O’Connor P.M., 2014. The Earliest Colubroid-Dominated Snake Fauna from Africa: Perspectives from the Late Oligocene Nsungwe Formation of Southwestern Tanzania. PLoS ONE, 9(3): e90415. doi: 10.1371/journal.pone.0090415.
    Orliac M.J., Ladevèze S., Gingerich P.D., Lebrun R. & Smith T., 2014. Endocranial morphology of Palaeocene Plesiadapis tricuspidens and evolution of the early primate brain. Proceedings of the Royal Society B, 281: 2013-2792. doi: 10.1098/rspb.2013.2792.
    Smith T., Quesnel F., De Plöeg G., De Franceschi D., Métais G., De Bast E., Solé F., Folie A., Boura A., Claude J., Dupuis C., Gagnaison C., Iakovleva A., Martin J., Maubert F., Prieur J., Roche E., Storme J.-Y., Thomas R., Tong H., Yans J. & Buffetaut E., 2014. First Clarkforkian Equivalent Land Mammal Age in the Latest Paleocene Basal Sparnacian Facies of Europe: Fauna, Flora, Paleoenvironment and (Bio)stratigraphy. PLoS ONE, 9(1): e86229. doi: 10.1371/journal.pone.0086229.
    Kocsis L., Gheerbrant E., Mouflih M., Cappetta H., Yans J. & Amaghzaz M., 2014. Comprehensive stable isotope investigation of marine biogenic apatite from the late Cretaceous-early Eocene phosphate Series of Morocco. Palaeogeography, Palaeoclimatology, Palaeoecology, 394: 74-88. doi: 10.1016/j.palaeo.2013.11.002.